From the Scottish Natural Capital Accounts report of the Office for National Statistics:

In 2015, the partial asset value of Scottish natural capital was estimated to be £273 billion, 34% of the UK asset valuation.

A quarter of the asset value was attributable to non-material benefits not directly captured in gross domestic product.

https://www.ons.gov.uk/economy/nationalaccounts/uksectoraccounts/bulletins/scottishnaturalcapital/ecosystemserviceaccounts2019#main-points